Robbie Henshaw will partner Bundee Aki at Center of the World Cup in Japan at Irelands Pool A decider against Samoa on Saturday.
The British and Irish Lions powerhouse has missed all three of Irelands World Cup games so far, in training later landing in Japan, with picked up a hamstring injury.
However, now that the 26-year-old is ready and fit to fire into a match Ireland must win with a bonus point to ensure their place.
Ireland have rolled out the bulk of their large guns to have the Samoans, however Jordan Larmour has set in at full-back in place of Rob Kearney.
Tadhg Beirne replaces his Munster colleague Peter OMahony in blindside flanker, with manager Joe Schmidt keen to ease the workload on the Irish and British Lions back-rower.
While Joey Carbery carries a seat on the bench, together with Ireland buoyed to have back to full 26, johnny Sexton begins at fly-half .
Ireland: 15 Jordan Larmour, 14 Keith Earls, 13 Robbie Henshaw,12 Bundee Aki, 11 Jacob Stockdale, 10 Jonathan Sexton, 9 Conor Murray, 1 Cian Healy, 2 Rory Best (c), 3 Tadhg Furlong, 4 Iain Henderson, 5 James Ryan, 6 Tadhg Beirne, seven Josh van der Flier, 8 CJ Stander
Replacements: 23 Andrew Conway, 16 Niall Scannell, 17 Dave Kilcoyne, 18 Andrew Porter, 19 Jean Kleyn, 20 Peter OMahony, 21 Luke McGrath, 22 Joey Carbery
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